BEAR DOG

Amphicyonidae is an extinct family of large terrestrial carnivores belonging to the suborder Caniformia (meaning "dog-like") and which inhabited North America, Europe, Asia, and Africa from the Middle Eocene subepoch to the Pleistocene epoch 46.2—1.8 Mya, existing for approximately  million years.Amphicyonids, often referred to as "bear 

dogs", crossed from Europe to North America during the Miocene epoch and are considered an Old World taxon. The earliest to appear is the (rather large)Ysengrinia (30—20 Mya), followed by Cynelos (24—7 Mya) and Amphicyon (23—5 Mya). These animals would have followed ungulates and other mammals to the New World for a period of approximately 7 million years. The New World amphicyonids of the subfamilies Daphoeninae (42-16 Mya) and Temnocyoninae (33-20 Mya) coexisted with the Old World counterparts. Note that the (often similar looking) members of the family Hemicyonidae are often called "bear-dogs" as well (although they are increasingly referred to as "dog-bears" to avoid confusion).While amphicyonids have traditionally been viewed as closely related to ursids (bears), some evidence suggests that they may instead be basal caniforms. (Hunt, 2004b). They were about as tall as the American black bear and were most likely ambushers because their legs were made for short, sudden bursts of speed. Bear-dog also nested their young in underground burrows.During the early Miocene, a number of large amphicyonids migrated from Eurasia into North America. These taxa belong to the Old World amphicyonid sub-family Amphicyoninae. The earliest to appear is the large bear dog Ysengrinia Ginsburg, followed by Cynelos Jourdan, and then by Amphicyon. This influx of amphicyonines, accompanied by other Old World ungulates and small mammals, indicates a prolonged interval (from 23 to 16.5 Ma) of faunal exchange between Asia and North America in the early Miocene, using the trans-Beringianroute.New World daphoenines (Daphoenodon, Borocyon) and temnocyonines coexisted with Old World amphicyonines
 (Ysengrinia, Amphicyon, Cynelos) 23.7-17.5 million years ago. These are the largest terrestrial carnivorans 50 kilograms (110 lb) to 200 kilograms (440 lb) that evolved on the North American continent up to this time. The immigrant 

amphicyoninesYsengrinia, Cynelos and Amphicyon appear at 23, 19.2, and 18.8 Ma, respectively, and herald the beginning of a Eurasian amphicyonine migration into North America that continued into the mid-iocene.Amphicyonids were as small as 5 kilograms (11 lb) and as large as 100 to 600 kilograms (220 to 1,300 lb) and evolved from wolf-like to bear-like.The diet of the amphicyonids was fully carnivorous as opposed to hypercarnivorous to mesocarnivorous in Canidae.

PTERANODON: THE HOODED MONSTER

Pteranodon from the Late Cretaceous geological period of North America in present day Kansas, Alabama, Nebraska, Wyoming, and South Dakota, was one of the largest pterosaur genera and had a maximum wingspan of over 6 metres (20 ft). Pteranodon is known from more fossil specimens than any other pterosaur, with about 1,200 specimens known to science, many of them well preserved, with complete skulls and articulated skeletons. It was an important genus of the animal community present in the Western Interior Seaway.Adult Pteranodon specimens from both major species can be divided into two distinct size classes. The smaller class of specimens have small, rounded head crests and very wide pelvic canals, even wider than those of the much larger size class. The size of the pelvic canal was probably to allow the laying of eggs, indicating that these smaller adults are females. The larger size class, representing male individuals, have narrow hips and very large crests, which were probably for display.Pteranodon was a reptile, but not a dinosaur. By definition, all dinosaurs belong to the groups Saurischia and Ornithischia, which excludes pterosaurs. Nevertheless, Pteranodon is frequently featured in dinosaur books and is strongly associated with dinosaurs by the general public.Specimens assigned to Pteranodon have been found in both the Smoky Hill Chalk deposits of the Niobrara Formation, and the slightly younger Sharon Springs deposits of the Pierre Shale Formation. WhenPteranodon was alive, this area was covered by a large inland sea, known as the Western Interior Seaway. Famous for fossils collected since 1870, these formations extend from as far south as Kansas in theUnited States to Manitoba in Canada. However, Pteranodon specimens (or any pterosaur specimens) have only been found in the southern half of the formation, in Kansas, Wyoming, and South Dakota. Despite the fact that numerous fossils have been found in the contemporary parts of the formation in Canada, no pterosaur specimens have ever been found there. This strongly suggests that the natural geographic range of Pteranodon covered only the southern part of the Niobrara, and that it's habitat did not extend farther north than South Dakota.

Some very fragmentary fossils belonging to pteranodontian pterosaurs, and possibly Pteranodon itself, have also been found on the Gulf Coast and East Coast of the United States. For example, some bone fragments from the Mooreville Formation of Alabama and the Merchantville Formation of Delaware may have come from Pteranodon, though they are too incomplete to make a definite identification.Some remains from Japan have also been tentatively attributed to Pteranodon, but their distance from its known Western Interior Seaway habitat makes this identification unlikelyAdult male Pteranodon were among the largest pterosaurs, and were the largest flying reptiles known until the late 20th Century, when the giant azhdarchid pterosaurs were discovered. The wingspan of an average adult male Pteranodon was 5.6 metres (18 ft). Adult females were much smaller, averaging 3.8 metres (12 ft) in wingspan. The largest specimen of Pteranodon longiceps from the Niobrara Formation measured 6.25 metres (20.5 ft) from wingtip to wingtip. An even larger specimen is known from the Pierre Shale Formation, with a wingspan of 7.25 metres (23.8 ft), though this specimen may belong to the distinct genus and species Geosternbergia maysei.While most specimens are found crushed, enough fossils exist to put together a detailed description of the animal.Pteranodon longiceps would have shared the sky with the giant-crested pterosaur Nyctosaurus. Compared to P. longiceps, which was a very common species,Nyctosaurus was rare, making up only 3% of pterosaur fossils from the formation. Also less common was the early toothed bird, Ichthyornis.

Methods used to estimate the weight of large male Pteranodon specimens (those with wingspans of about 7 meters) have been notoriously unreliable, producing a wide range of estimates from as low as 20 kilograms (44 lb) and as high as 93 kilograms (210 lb). In a review of pterosaur size estimates published in 2010, researchers Mark Witton and Mike Habib demonstrated that the latter, largest estimates are almost certainly incorrect given the total volume of a Pteranodon body, and could only be correct if the animal "was principally comprised of aluminium." Witton and Habib considered the methods used by researchers who obtained smaller weight estimates equally flawed. Most have been produced by scaling modern animals such as bats and birds up to Pteranodon size, despite the fact that pterosaurs have vastly different body proportions and soft tissue anatomy than any living animal.It is likely that, as in other polygynous animals (in which males compet for association with harems of females), Pteranodon lived primarily on offshore rookeries, where they could nest away from land-based predators and feed far from shore; most Pteranodon fossils are found in locations which at the time, were hundreds of kilometres from the coastline.Below the surface, the sea was populated primarily by invertebrates such as ammonites and squid. Vertebrate life, apart from basal fish, included sea turtles such asToxochelys, the plesiosaur Styxosaurus, and the flightless diving bird Parahesperornis. Mosasaurs were the most common marine reptiles, with genera including Clidastes and Tylosaurus. At least some of these marine reptiles are known to have fed on Pteranodon. Barnum Brown, in 1904, reported plesiosaur stomach contents containing "pterodactyl" bones, most likely from Pteranodon.The diet of Pteranodon is known to have included fish; fossilized fish bones have been found in the stomach area of one Pteranodon, and a fossilized fish bolus has been found between the jaws of another Pteranodon, specimen AMNH 5098. Numerous other specimens also preserve fragments of fish scales and vertebrae near the torso, indicating that fish made up a majority of the diet of Pteranodon (though they may also have taken invertebrates).Fossils from terrestrial dinosaurs also have been found in the Niobrara Chalk, suggesting that animals who died on shore must have been washed out to sea (one specimen of a hadrosaur appears to have been scavenged by a shark).Traditionally, most researchers have suggested that Pteranodon would have taken fish by dipping their beaks into the water while in low, soaring flight. However, this was probably based on the assumption that the animals could not take off from the water surface.It is more likely that Pteranodon could take off from the water, and would have dipped for fish while swimming rather than while flying. Even a small, female Pteranodon could have reached a depth of at least 80 centimetres (31 in) with its long bill and neck while floating on the surface, and they may have reached even greater depths by plunge-diving into the water from the air like some modern long-winged seabirds. In 1994, Bennett noted that the head, neck, and shoulders of Pteranodon were as heavily built as diving birds, and suggested that they could dive by folding back their wings like the modern Gannet.

QUETZACOATLUS :THE KING OF HEAVEN

Quetzalcoatlus was a pterodactyloid pterosaur known from the Late Cretaceous of North America (Maastrichtian stage, about 68–65.5 million years ago), and one of the largest known flying animals of all time. 

It was a member of the Azhdarchidae, a family of advanced toothless pterosaurs with unusually long, stiffened necks. Its name comes from the Mesoamerican feathered serpent godQuetzalcoatl.During the 19th century, in England many fragmentary pterosaur fossils were found in the Cambridge Greensand, a layer from the early Cretaceous, that had originated as a sandy seabed. Decomposing pterosaur cadavers, floating on the sea surface, had gradually lost individual bones that sank to the bottom of the sea. Water currents then moved the bones around, eroding and polishing them, until they were at last covered by more sand and fossilised. Even the largest of these remains were damaged and difficult to interpret. They had been assigned to the genus Pterodactylus, as was common for any pterosaur species described in the early and middle 19th century.Young researcher Harry Govier Seeley was commissioned to bring order to the pterosaur collection of the Sedgwick Museum in Cambridge. He soon concluded that it was best to create a new genus for the Cambridge Greensand material that he named Ornithocheirus, "bird hand", as he in this period still considered pterosaurs to be the direct ancestors of birds, and assumed the hand of the genus to represent a transitional stage in the evolution towards the bird hand. To distinguish the best pieces in the collection, and partly because they had already been described as species by other scientists, he in 1869 and 1870 each gave them a separate species name: O. simus, O. woodwardi, O. oxyrhinus, O. carteri, O. platyrhinus, O. sedgwickii, O. crassidens, O. capito, O. eurygnathus, O. reedi, O. cuvieri, O. scaphorhynchus, O. brachyrhinus, O. colorhinus, O. dentatus, O. denticulatus, O. enchorhynchus, O. 

xyphorhynchus, O. fittoni, O. nasutus, O. polyodon, O. compressirostris, O. tenuirostris, O. machaerorhynchus, O. platystomus, O. microdon, O. oweni and O. huxleyi, thus 28 in total. As yet Seeley did not designate atype species.When Seeley published his conclusions in his 1870 book The Ornithosauria, this provoked a reaction by the leading British paleontologist of his day, Richard Owen. Owen was not an evolutionist and he therefore considered the name Ornithocheirus to be inappropriate; he also thought it was possible to distinguish two main types within the material, based on differences in snout form and tooth position — the best fossils consisted of jaw fragments. He in 1874 created two new genera: Coloborhynchus and Criorhynchus. Coloborhynchus, "maimed beak", comprised a new species, Coloborhynchus clavirostris, the type species, and two species reassigned fromOrnithocheirus: C. sedgwickii and C. cuvieri. Criorhynchus, "ram beak", consisted entirely of former Ornithocheirus species: the type species Criorhynchus simus and furthermore C. eurygnathus, C. capito, C. platystomus, C. crassidens and C. reedi.In 1914 Reginald Walter Hooley made a new attempt to structure the large number of species. Keeping the name Ornithocheirus, he added to it Owen's Criorhynchus, in which however Coloborhynchus was sunk, and to allow for a greater differentiation created two new genera, again based on jaw form: Lonchodectes and Amblydectes. Lonchodectes, "lance biter", comprised L. compressirostris, L. giganteus and L. daviesii. Amblydectes, "blunt biter", consisted of A. platystomus, A. crassidens and A. eurygnathus. However, Hooley's classification was rarely applied later in the century, when it became common to subsume all the poorly preserved and confusing material under the name Ornithocheirus. In 1978 Peter Wellnhofer, assuming no type species had been designated, made Ornithocheirus compressirostris the type.Seeley did not accept Owen's position. In 1881 he designated O. simus the type species of Ornithocheirus and named a new species O. bunzeli. In 1888 Edward Newton renamed several existing species names into: Ornithocheirus clavirostris, O. daviesii, O. sagittirostris, O. validus and O. giganteus; as new species he created: O. clifti, O. diomedeus, O. nobilis and O. curtus. Others had already named an O. umbrosus, O. harpyia, O. macrorhinus and O. hilsensis and would create an O. hlavaci,O. wiedenrothi and O. mesembrinus.From the seventies onwards many new pterosaur fossils were found in Brazil, from formations about the same age as the Cambridge Greensand, 110 million years old. Contrary to the English material, these new finds included some of the best preserved large pterosaur skeletons and several new genera names were given to them, such as Anhanguera. This situation caused a renewed interest in the Ornithocheirusmaterial and the validity of the several names based on it, for it might be possible that it could by more detailed studies be established that the Brazilian pterosaurs were actually junior synonyms of the European types. Several European researchers concluded that this was indeed the case. Unwin revived Coloborhynchus and Michael Fastnacht Criorhynchus, each author ascribing Brazilian species to these genera. However, in 2000 Unwin stated that Criorhynchus could not be valid. Referring to Seeley's designation of 1881 he considered Ornithocheirus simus, holotype CAMSM B.54428, to be the type species. This also made it possible to revive Lonchodectes, using as type the former O. compressirostris, which then became L. compressirostris. This position has not universally been accepted. Brazilian workers also typically reject the identification of their genera with European types. Unwin, and this caused no controversy, reaffirmed that most Ornithocheirus species are nomina dubia, names that are invalid because the fossils they refer to lack sufficient diagnostic features.As a result, though over forty species have been named in the genus Ornithocheirus over the years, not a single one of them, not even O. simus, is currently recognized as valid by all pterosaur researchers. Often, there is a total lack of consensus; e.g.Tropeognathus mesembrinus named by Peter Wellnhofer in 1987 has afterwards been considered Ornithocheirus mesembrinus by David Unwin in 2003 (making Tropeognathus a junior synonym) , but as Anhanguera mesembrinus by Alexander Kellner in 1989, Coloborhynchus mesembrinus by André Veldmeijer in 1998 and Criorhynchus mesembrinus by Michael Fastnacht in 2001. Even earlier, in 2001, Unwin had referred the "Tropeognathus" material to O. simus in which he was followed by Veldmeijer; however the latter, denying that O. simus is the type species instead of O. compressirostris, now uses the name Criorhynchus simus. Kellner in 2000 again recognized Tropeognathus as a valid genus.

TERATORNIS: THE DEATH BIRD

Teratornis merriami (Merriam's Teratorn) was a huge North American teratorn, with a wingspan of around 3.5 to 3.8 meters (11 to 12 feet) and a wing area of 17.5 square meters, standing an estimated 75 cm tall and weighing about 15 kg. It was somewhat larger than the extant Andean Condor and nearly two times as heavy as the California Condor. A closely related genus, Aiolornis, was about 40% larger and lived at an earlier time; it was formerly known 

as Teratornis incredibilis, but is distinct enough to be placed in its own genus.T. merriami is the best-known of the teratorns. A large number of fossil and subfossil bones, representing more than 100 individuals, have been found in locations in California, southern Nevada, Arizona, and Florida, though most are from the Californian La Brea Tar Pits. All remains but one Early Pleistocenepartial skeleton from the Leisey Shell Pit near Charlotte Harbor, Florida (which may represent a different species or a subspecies) date from the Late Pleistocene, with the youngest remains dating from the Pleistocene-Holocene boundary.The finger bones are fused as in all modern birds; however, part of the index finger forms a shelf which aided in bearing the load of long and stout primaries, which enabled the bird to utilize strong upcurrents. The legs were similar to an Andean Condor's, but stouter, and the feet were able to hold prey items for tearing off pieces, but not able to exert a very forceful grip such as in birds of prey. Its wing loading was not much larger than a Californian Condor's, and Merriam's Teratorn should have been able to take off by simply jumping and beating its wings under most circumstances. (Campbell & Tonni, 1983). Indeed, it seems to have been better adapted for that than for utilizing a short run into the wind from an elevated location as condors do, as its legs are proportionally smaller and its stride less than in condors (Fisher, 1945).T. merriami generally lived in a manner similar to condors, although its larger bill suggests that it was a more active predator. Prey up to the size of a small rabbit would probably have been swallowed more or less whole, while carrion would have been fed on in a manner similar to that of condors or vultures. The large number of finds in the La Brea Tar Pits were usually considered to be from teratorns which were attracted by Pleistocene megafauna that became stuck in the viscous asphalt trying to drink from pools of water that gathered on the surface and died, with the teratorns subsequently falling victim to the sticky deposits too. Merriam's Teratorn probably played an important role in opening up the body cavities of carcasses for smaller birds like eagles and ravens which are also known to have frequented the locality, as mammalian predators, being unable to fly, could hardly reach most carcasses without getting mired in the asphalt themselves.

TITANOBOA: THE SNAKE KING

The world's biggest snake was a massive anaconda-like beast that slithered through steamy tropical rain forests about 60 million years ago, says a new study that describes the ancient giant.

Fossils found in northeastern Colombia's Cerrejon coal mine indicate the reptile, dubbed Titanoboa cerrejonesis, was at least 42 feet (13 meters) long and weighed 2,500 pounds (1,135 kilograms).

"That's longer than a city bus and … heavier than a car," said lead study author Jason Head, a

 fossil-snake expert at the University of Toronto Mississauga in Canada and a research associate with the Smithsonian Institution.Previously the biggest snake known was Gigantophis garstini, which was 36 to 38 feet (11 to 11.6 meters) long. That snake lived in North Africa about 40 million years ago.

Hans-Dieter Sues, associate director for research and collections at the Smithsonian's National Museum of Natural History, was not involved with  the study but has seen the snake fossils.

Sues noted that humans would stand no chance against one of these giants, which killed their prey by slow suffocation."Given the sheer size—the sheer cross-section of that snake—it would be probably like one of those devices they use to crush old cars in a junkyard," Sues said.In addition, the snake's heft indicates that it lived when the tropics were much warmer than they are today, a find that holds potential implications for theories of once and future climate change.Biggest Snake Needed the HeatScientists know there's a link between a snake's body size, how fast it uses and produces energy, and climate.(Related: "World's Smallest Snake Discovered, Study Says" [August 3, 20008].)"We were able to use the snake, if you will, as a giant fossil thermometer," study author Head said.His team found that, for Titanoboa to reach its epic proportions,  mean year-round temperatures would have been about 90 degrees Fahrenheit (32 degrees Celsius)—significantly hotter than today's tropics.This supports the idea that tropical temperatures spike as the rest of the world heats up due to global warming, the study authors say.The competing theory is that, during bouts of warming, the tropics stay about the same average temperatures as they are today while areas north and south of the Equator heat up.James Zachos, an expert on ancient climates at the University of California, Santa Cruz, who was not involved in the study, agreed.As the biggest known snake, Titanoboa supports the idea of "much hotter tropics during extreme greenhouse periods," Zachos said.Big Reptiles on the Horizon?Study co-author Jonathan Bloch is a vertebrate paleontologist at the University of Florida's Museum of Natural History in Gainesville.The same Colombian coal mine that contained the biggest snake also yielded massive turtles and crocodiles, he said."You can think about it as an ecosystem dominated by giants, I think, and these are probably giants that got large because of the warmer mean annual temperature," he said.The findings, detailed in this week's issue of the journal Nature, paint a picture of what the future might hold if supercharged global warming takes place.According to some models, global temperatures could approach the same levels that gave rise to the biggest snake by the end of this century.If current greenhouse gas emissions continue apace, there's a chance snakes the size of Titanoboa could return, Bloch said."Or maybe snakes would go extinct in the tropics," he said. "In other words, the warming could happen so rapidly that they wouldn't have time to adapt.

PERMIAN EXTICION: THE GREAT DYING

The Permian extinction event, informally known as the Great Dying, was an extinction event that occurred 251.4 Ma (million years ago), forming the boundary between the Permian and Triassic geologic periods. It was the Earth's most severe extinction event, with up to 96% of all marine speciesand 70% of terrestrial vertebrate species becoming extinct  It is the only known 

mass extinction of insects.Some 57% of all families and 83% of all genera were killed. Because so much biodiversity was lost, the recovery of life on Earth took significantly longer than after other extinction events.This event has been described as the "mother of all mass extinctions."Researchers have variously suggested that there were from one to three distinct pulses, or phases, of extinction.There are several proposed mechanisms for the extinctions; the earlier phase was likely due to gradual environmental change, while the latter phase has been argued to be due to a catastrophic event. Suggested mechanisms for the latter include large or multiple bolide impact events, increased volcanism, and sudden release of methane clathrate from the sea floor; gradual changes include sea-level change, anoxia, increasing aridity, and a shift in ocean circulation driven by climate change.There are several proposed mechanisms for the extinction event, including both catastrophic and gradualistic processes (similar to those theorized for the Cretaceous–Tertiary extinction event). The former include large or multiple bolide impact events, increased volcanism, or sudden release of methane hydrates from the sea floor. The latter include sea-level change,anoxia, and increasing aridity. Any hypothesis about the cause must explain the selectivity of the event, which primarily affected organisms with calcium carbonate skeletons; the long (4–6 million year) period before recovery started; and the minimal extent of biological mineralization (despite inorganic carbonates being deposited) once the recovery began.

KAPROSUCHUS

Kaprosuchus is an extinct genus of mahajangasuchid crocodyliform. It is known from a single nearly complete skull collected from theUpper Cretaceous Echkar Formation of Niger. The name means "boar crocodile" from the Greek kapros ("boar") and souchos ("crocodile") in reference to its unusually large caniniform teeth which resemble those of 

a boar.It has been nicknamed "BoarCroc" by Paul Serenoand Hans Larsson, who first described the genus in a monograph published in ZooKeys in 2009 along with other Saharan crocodyliformes such as Anatosuchus and Laganosuchus. The type species is K. saharicus.Kaprosuchus is estimated to have been around 6 metres (20 feet) in length. It possesses three sets of tusk-like caniniform teeth that project above and below the skull, one of which in the lower jaw fits into notches in upper jaw. This type of dentition is not seen in any other known crocodyliform. Another unique characteristic of Kaprosuchus is the presence of large, rugose horns formed from the squamosal and parietal bones that project posteriorly from the skull. Smaller projections are also seen in the closely related Mahajangasuchus.The snout of Kaprosuchus shows generalized proportions and the naris is positioned dorsally. In Kaprosuchus many teeth are hypertrophied and labiolingually (laterally) compressed, unlike those of crocodyliforms with similarly shallow snouts, which are usually subconical and of moderate length. Another difference between the skull of Kaprosuchus and those of crocodyliforms that also possess dorsoventrally compressed snouts is the great depth of the posterior portion of the skull.In Kaprosuchus, the orbits (i.e., eye sockets) open laterally and are angled slightly forward rather than upward. The orbits turned forward suggest that there was somewhat stereoscopic vision, i.e., an overlap in the visual field of the animal.The surfaces of the premaxillae are rugose with the edges elevated above the body of the bone, suggesting that a keratinous shield would have been supported by the rugosities at the tip of the snout. Along the interpremaxillary suture, the area where the two premaxillae meet, the surface is smooth, giving the paired rugosity of the premaxillae the resemblance of a moustache in anterior view.

PANGEA: BIGGER THAN EVER

Pangea was the supercontinent that existed during the Paleozoic and Mesozoic eras about 250 million years ago, before the component continents were separated into their current configuration.The name was coined during a 1926 symposium discussing Alfred Wegener's theory of continental drift. In his book The Origin of Continents and Oceans (Die Entstehung der Kontinente und Ozeane) first published in 1915, he postulated that all the continents had at one time formed a single supercontinent which he called the "Urkontinent", before later breaking 

up and drifting to their present locations.The single enormous ocean which surrounded Pangaea was accordingly named Panthalassa.There were three major phases in the break-up of Pangaea. The first phase began in the Early-Middle Jurassic (about 175 Ma), when Pangaea began to rift from the Tethys Ocean in the east and the Pacific in the west, ultimately giving rise to the supercontinents Laurasia and Gondwana. The rifting that took place between North America and Africa produced multiplefailed rifts. One rift resulted in a new ocean, the North Atlantic Ocean.The Atlantic Ocean did not open uniformly; rifting began in the north-central Atlantic. The South Atlantic did not open until the Cretaceous. Laurasia started to rotate clockwise and moved northward with North America to the north, and Eurasia to the south. The clockwise motion of Laurasia also led to the closing of the Tethys Ocean. Meanwhile, on the other side of Africa, new rifts were also forming along the adjacent margins of east Africa, Antarctica and Madagascar that would lead to the formation of the southwestern Indian Ocean that would also open up in the Cretaceous.The second major phase in the break-up of Pangaea began in the Early Cretaceous (150–140 Ma), when the minor supercontinent of Gondwana separated into multiple continents (Africa, South America, India, Antarctica, and Australia). About 200 Ma, the continent of Cimmeria, as mentioned above (see "Formation of Pangaea"), collided with Eurasia. However, a subduction zone was forming, as soon as Cimmeria collided.This subduction zone was called the Tethyan Trench. This trench might have subducted what is called the Tethyan mid-ocean ridge, a ridge responsible for the Tethys Ocean's expansion. It probably caused Africa, India and Australia to move northward. In the Early Cretaceous, Atlantica, today's South America and Africa, finally separated from eastern Gondwana (Antarctica, India and Australia), causing the opening of a "South Indian Ocean". In the Middle Cretaceous, Gondwana fragmented to open up the South Atlantic Ocean as South America started to move westward away from Africa. The South Atlantic did not develop uniformly; rather, it rifted from 

south to north.Also, at the same time, Madagascar and India began to separate from Antarctica and moved northward, opening up the Indian Ocean. Madagascar and India separated from each other 100–90 Ma in the Late Cretaceous. India continued to move northward toward Eurasia at 15 centimeters (6 in) per year (a plate tectonic record), closing the Tethys Ocean, while Madagascar stopped and became locked to the African Plate. New Zealand, New Caledonia and the rest of Zealandia began to separate from Australia, moving eastward towards the Pacific and opening the Coral Sea and Tasman Sea.The third major and final phase of the break-up of Pangaea occurred in the early Cenozoic (Paleocene to Oligocene). Laurasia split when North America/Greenland (also called Laurentia) broke free from Eurasia, opening the Norwegian Sea about 60–55 Ma. The Atlantic and Indian Oceans continued to expand, closing the Tethys Ocean.Meanwhile, Australia split from Antarctica and moved rapidly northward, just as India did more than 40 million years earlier, and is currently on a collision course with eastern Asia. Both Australia and India are currently moving in a northeastern direction at 5–6 centimeters (2–3 in) per year. Antarctica has been near or at the South Pole since the formation of Pangaea about 280 Ma. India started to collide with Asia beginning about 35 Ma, forming the Himalayan orogeny, and also finally closing the Tethys Seaway; this collision continues today. The African Plate started to change directions, from west to northwest toward Europe, and South America began to move in a northward direction, separating it from Antarctica and allowing complete oceanic circulation around Antarctica for the first time. The latter of which, together with decreasing atmospheric carbon dioxide concentrations caused a rapid cooling of Antarctica and allowed glaciers to form, which eventually coalesced into the kilometers thick ice sheets we see today. Other major events took place during the Cenozoic, including the opening of the Gulf of California, the uplift of the Alps, and the opening of the Sea of Japan. The break-up of Pangaea continues today in the Great Rift Valley.

SYNAPSIDS

Synapsids ('fused arch') are a group of animals that includes mammals and everything more closely related to mammals than to other livingamniotes. They are easily separated from other amniotes by having an opening low 
 in the skull roof behind each eye, leaving a bony archbeneath each, accounting for their name.Primitive synapsids are usually called pelycosaurs; more advanced mammal-like ones, therapsids. The non-mammalian members are described as mammal-like reptiles in classical systematics, but are referred to as "stem-mammals" or "proto-mammals" under cladistic terminology. Synapsids evolved from basal amniotes and are one of the two major groups of the later amniotes, the other major group being the sauropsids (reptiles and birds). They are distinguished from other amniotes by having a single opening (temporal fenestra) in their skull behind each eye, which developed in the ancestral synapsid about 324 million years ago (mya) during the late Carboniferous Period.

Synapsids were the dominant terrestrial animals in the middle to late Permian period. As with almost all groups then extant, their numbers and variety were severely reduced by the Permian extinction. Some species survived into the Triassic period, but archosaurs quickly became the dominant animals and few of the non-mammalian synapsids outlasted the Triassic, although survivors persisted into the Cretaceous. However, as a phylogenetic unit they included the mammal descendants, and in this sense synapsids are still very much a living group of vertebrates. In the form of mammals, Synapsids (most recently and notably humans) again became the dominant land animals after they outcompeted birds following the K-T extinction event.

THE PENKNIFE

There is currently disagreement about the function of the enlarged "sickle claw" on the second toe. When John Ostrom described it for Deinonychus in 1969, he interpreted the claw as a blade-like slashing weapon, much like the canines of some saber-toothed cats, used with powerful kicks to cut into prey. Adams (1987) suggested that the talon was used to disembowel large ceratopsian dinosaurs. The interpretation of the sickle claw as a killing weapon applied to all dromaeosaurids. However, Manning et al. argued that the claw instead served as a hook, reconstructing the keratinous sheath with an elliptical cross section, instead of the previously inferred inverted teardrop shape. In Manning's interpretation, the second toe claw would be used as a climbing aid when subduing bigger prey and also as stabbing weapon.

Ostrom compared Deinonychus to the ostrich and cassowary. He noted that the bird species can inflict serious injury with the large claw on the second toe. The cassowary has claws up to 125 millimetres (4.9 in) long. Ostrom cited Gilliard (1958) in saying that they can sever an arm or disembowel a man.Kofron (1999 and 2003) studied 241 documented cassowary attacks and found that one human and two dogs had been killed, but no evidence that cassowaries can disembowel or dismember other animals^. Cassowaries use their claws to defend themselves, to attack threatening animals, and in agonistic displays such as the Bowed Threat Display. The seriema also has an enlarged second toe claw, and uses it to tear apart small prey items for swallowing.
Phillip Manning and colleagues (2009) attempted to test the function of the sickle claw and similarly shaped claws on the forelimbs. They analyzed the bio-mechanics of how stresses and strains would be distributed along the claws and into the limbs, using X-ray imaging to create a three dimensional contour map of a forelimb claw from Velociraptor. For comparison, they analyzed the construction of a claw from a modern predatory bird, the Eagle Owl. They found that, based on the way that stress was conducted along the claw, they were ideal for climbing. The scientists found that the sharpened tip of the claw was a puncturing and gripping instrument, while the curved and expanded claw base helped transfer stress loads evenly.
The Manning team also compared the curvature of the dromaeosarid "sickle claw" on the foot with curvature in modern birds and mammals. Previous studies had shown that the amount of curvature in a claw corresponded to what lifestyle the animal has: animals with strongly curved claws of a certain shape tend to be climbers, while straighter claws indicate ground-dwelling lifestyles. The sickle-claws of the dromaeosaurid Deinonychus have a curvature of 160 degrees, well within the range of climbing animals. The forelimb claws they studied also fell within the climbing range of curvature.
Paleontologist Peter Mackovicky commented on the Manning team's study, stating that small, primitive dromaeosaurids (such as Microraptor) were likely to have been tree-climbers, but that climbing did not explain why later, gigantic dromaeosaurids such as Achillobator retained highly curved claws when they were too large to have climbed trees. Mackovickey speculated that giant dromaeosaurids may have adapted the claw to be used exclusively for latching on to prey.

SPINOSAUR: THE LEGEND OF A KILLER

A recent movie battle pitted a Spinosaurus against Tyrannosaurus rex, with the former portrayed as a victor after it snapped its rival’s neck. While Spinosaurus and T. rex never fought in real life, since they lived millions of miles and thousands of years apart, Spinosaurus holds the world record for being the largest known carnivorous dinosaur, given its impressive length and 9.9-ton build.

Sail
The imposing dinosaur’s most unusual feature was its large sail. Whenever Spinosaurus would arch its back, the sail, made of lengthy spines covered with skin, would rise into the air. The sail alone was the height of a human basketball star. Paleontologists continue to debate its function, but most suspect the sail helped to regulate body temperature and was used to woo the opposite sex or to scare off competing males.
Teeth and Diet
It’s no wonder Spinosaurus often makes film appearances, since this dinosaur flashed a million-dollar "smile." While most carnivorous dinos had curved teeth, the teeth of Spinosaurus were straight and probably functioned like knives, skewering often-slippery prey. Like modern grizzly bears, Spinosaurus probably spent a lot of time grabbing fish in, or near, water. A dinosaur of such girth, however, likely did not subsist on an all seafood diet. It probably killed and consumed smaller dinosaurs too, along with scavenging flesh from corpses.
Big Head
Such feasting would not have posed a challenge, since Spinosaurus possessed what was arguably the longest head of any known carnivorous dinosaur. Measuring close to 6 feet in length, the head featured a narrow snout — all the better for showcasing its straight teeth — with tiny ears on either side. Scaly skin covered its neck.

THE FIGHTING DINOSAURS

The "Fighting Dinosaurs" specimen, found in 1971, preserves a Velociraptor mongoliensis and Protoceratops andrewsi in combat and provides direct evidence of predatory behavior. When originally reported, it was hypothesized that the two animals drowned. However, as the animals were preserved in ancient sand dune deposits, it is now thought that the animals were buried in sand, either from a collapsing dune or in a sandstorm. Burial must have been extremely fast, judging from the lifelike poses in which the animals were preserved. Both forelimbs and one hindlimb of the

 Protoceratops are missing, which has been seen as evidence of scavenging by other animals.The distinctive claw, on the second digit of dromaeosaurids, has traditionally been depicted as a slashing weapon; its assumed use being to cut and disembowel prey. In the "Fighting Dinosaurs" specimen, the Velociraptor lies underneath, with one of its sickle claws apparently embedded in the throat of its prey, while the beak of Protoceratops is clamped down upon the right forelimb of its attacker. This suggests Velociraptor may have used its sickle claw to pierce vital organs of the throat, such as the jugular vein, carotid artery, or trachea (windpipe), rather than slashing the abdomen. The inside edge of the claw was rounded and not unusually sharp, which may have precluded any sort of cutting or slashing action, although only the bony core of the claw is known. The thick abdominal wall of skin and muscle of large prey species would have been difficult to slash without a specialized cutting surface. The slashing hypothesis was tested during a 2005 BBC documentary, The Truth About Killer Dinosaurs. The producers of the program created an artificial Velociraptor leg with a sickle claw and used a pork belly to simulate the dinosaur's prey. Though the sickle claw did penetrate the abdominal wall, it was unable to tear it open, indicating that the claw was not used to disembowel prey. However, this experiment has not been published or repeated by other scientists, so its results cannot be confirmed.